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Genetic drivers of heterogeneity in type 2 diabetes pathophysiology

Nature 2024年第8003期627卷 347-357页

作者： Ken Suzuki Konstantinos Hatzikotoulas Lorraine Southam Henry J Taylor Xianyong Yin Kim M Lorenz Ravi Mandla Alicia Huerta-Chagoya Giorgio E M Melloni Stavroula Kanoni Nigel W Rayner Ozvan Bocher Ana Luiza Arruda Kyuto Sonehara Shinichi Namba Simon S K Lee Michael H Preuss Lauren E Petty Philip Schroeder Brett Vanderwerff Mart Kals Fiona Bragg Kuang Lin Xiuqing Guo Weihua Zhang Jie Yao Young Jin Kim Mariaelisa Graff Fumihiko Takeuchi Jana Nano Amel Lamri Masahiro Nakatochi Sanghoon Moon Robert A Scott James P Cook Jung-Jin Lee Ian Pan Daniel Taliun Esteban J Parra Jin-Fang Chai Lawrence F Bielak Yasuharu Tabara Yang Hai Gudmar Thorleifsson Niels Grarup Tamar Sofer Matthias Wuttke Chloé Sarnowski Christian Gieger Darryl Nousome Stella Trompet Soo-Heon Kwak Jirong Long Meng Sun Lin Tong Wei-Min Chen Suraj S Nongmaithem Raymond Noordam Victor J Y Lim Claudia H T Tam Yoonjung Yoonie Joo Chien-Hsiun Chen Laura M Raffield Bram Peter Prins Aude Nicolas Lisa R Yanek Guanjie Chen Jennifer A Brody Edmond Kabagambe Ping An Anny H Xiang Hyeok Sun Choi Brian E Cade Jingyi Tan K Alaine Broadaway Alice Williamson Zoha Kamali Jinrui Cui Manonanthini Thangam Linda S Adair Adebowale Adeyemo Carlos A Aguilar-Salinas Tarunveer S Ahluwalia Sonia S Anand Alain Bertoni Jette Bork-Jensen Ivan Brandslund Thomas A Buchanan Charles F Burant Adam S Butterworth Mickaël Canouil Juliana C N Chan Li-Ching Chang Miao-Li Chee Ji Chen Shyh-Huei Chen Yuan-Tsong Chen Zhengming Chen Lee-Ming Chuang Mary Cushman John Danesh Swapan K Das H Janaka de Silva George Dedoussis Latchezar Dimitrov Ayo P Doumatey Shufa Du Qing Duan Kai-Uwe Eckardt Leslie S Emery Daniel S Evans Michele K Evans Krista Fischer James S Floyd Ian Ford Oscar H Franco Timothy M Frayling Barry I Freedman Pauline Genter Hertzel C Gerstein Vilmantas Giedraitis Clicerio González-Villalpando Maria Elena González-Villalpando Penny Gordon-Larsen Myron Gross Lindsay A Guare Sophie Hackinger Liisa Hakaste Sohee Han Andrew T Hattersley Christian Herder Momoko Horikoshi Annie-Green Howard Willa Centre for Genetics and Genomics Versus Arthritis Centre for Musculoskeletal Research Division of Musculoskeletal and Dermatological Sciences University of Manchester Manchester UK. Department of Diabetes and Metabolic Diseases Graduate School of Medicine University of Tokyo Tokyo Japan. Department of Statistical Genetics Osaka University Graduate School of Medicine Suita Japan. Institute of Translational Genomics Helmholtz Zentrum München German Research Center for Environmental Health Neuherberg Germany. konstantinos.hatzikotoulas@helmholtz-munich.de. Institute of Translational Genomics Helmholtz Zentrum München German Research Center for Environmental Health Neuherberg Germany. Center for Precision Health Research National Human Genome Research Institute National Institutes of Health Bethesda MD USA. British Heart Foundation Cardiovascular Epidemiology Unit Department of Public Health and Primary Care University of Cambridge Cambridge UK. Heart and Lung Research Institute University of Cambridge Cambridge UK. Department of Epidemiology School of Public Health Nanjing Medical University Nanjing China. Department of Biostatistics and Center for Statistical Genetics University of Michigan Ann Arbor MI USA. Corporal Michael J. Crescenz VA Medical Center Philadelphia PA USA. Department of Systems Pharmacology and Translational Therapeutics University of Pennsylvania Perelman School of Medicine Philadelphia PA USA. Department of Genetics University of Pennsylvania Perelman School of Medicine Philadelphia PA USA. Programs in Metabolism and Medical and Population Genetics Broad Institute of Harvard and MIT Cambridge MA USA. Diabetes Unit and Center for Genomic Medicine Massachusetts General Hospital Boston MA USA. TIMI Study Group Division of Cardiovascular Medicine Brigham and Women's Hospital Harvard Medical School Boston MA USA. William Harvey Research Institute Barts and the London School of Medicine and Dentistry Queen Mary University of London L

Type 2 diabetes (T2D) is a heterogeneous disease that develops through diverse pathophysiological processes and molecular mechanisms that are often specific to cell type. Here, to characterize the genetic contribution to these processes across ancestry groups, we aggregate genome-wide association study data from 2,535,601 individuals (39.7% not of European ancestry), including 428,452 cases of T2D. We identify 1,289 independent association signals at genome-wide significance (P < 5 × 10) that map to 611 loci, of which 145 loci are, to our knowledge, previously unreported. We define eight non-overlapping clusters of T2D signals that are characterized by distinct profiles of cardiometabolic trait associations. These clusters are differentially enriched for cell-type-specific regions of open chromatin, including pancreatic islets, adipocytes, endothelial cells and enteroendocrine cells. We build cluster-specific partitioned polygenic scores in a further 279,552 individuals of diverse ancestry, including 30,288 cases of T2D, and test their association with T2D-related vascular outcomes. Cluster-specific partitioned polygenic scores are associated with coronary artery disease, peripheral artery disease and end-stage diabetic nephropathy across ancestry groups, highlighting the importance of obesity-related processes in the development of vascular outcomes. Our findings show the value of integrating multi-ancestry genome-wide association study data with single-cell epigenomics to disentangle the aetiological heterogeneity that drives the development and progression of T2D. This might offer a route to optimize global access to genetically informed diabetes care.

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Dynamical multiple polynomial-based neural networks classifier realized with the aid of dropfilter and dual statistical selection

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engineering Applications of Artificial Intelligence 2025年 157卷

作者： Zhen Wang Sung-Kwun Oh Zunwei Fu Seok-Beom Roh Witold Pedrycz Research Center for Big Data and Artificial Intelligence Linyi University Linyi 276005 China School of Automation and Electrical Engineering Linyi University Linyi 276005 Shandong China School of Electrical & Electronic Engineering The University of Suwon 17 Wauan-gil Bongdam-eup Hwaseong-si Gyeonggi-do 18323 South Korea Department of Electronic Engineering Seokyeong University Seoul 02713 South Korea Department of Computer Science The University of Suwon 17 Wauan-gil Bongdam-eup Hwaseong-si Gyeonggi-do 18323 South Korea Department of Electrical and Computer Engineering University of Alberta Edmonton AB T6G 2R3 Canada Systems Research Institute Polish Academy of Sciences 00-901 Warsaw Poland Department of Computer Engineering Faculty of Engineering and Natural Sciences Istinye University Sariyer Istanbul Turkiye

Polynomial neural networks (PNN) have emerged as an effective regression modeling methodology in computational intelligence, relying on its interpretable polynomial nodes to fit complex nonlinear data relationships and the adaptive nature of self-organizing networks. To break the bottleneck of PNN structure in the field of multi-classification, this study designs a dynamical multiple polynomial-based neural networks (DMPNN) classifier, focusing on developing a flexible polynomial network classification methodology that enhances predictive capabilities without sacrificing the advantages of PNN structures. Our approach effectively addresses the challenges of multi-class classification with uncertain class boundaries and reduces computational complexity, which is achieved through the synergy of several proposed techniques. Three key issues underpin the proposed DMPNN: (a) The integration of PNN regression models using the one-against-all strategy can provide effective and scalable solutions to multi-class classification problems, especially for uncertain class boundary issues. (b) The dual statistical selection (DSS) approach aims to eliminate redundant inputs during data processing, reduce the computational burden, and increase the variety of neural network nodes in the model neuron selection stage. (c) The synergy of regularization methods including the ℓ2 norm-based method (ℓ2-LSM) and the DropFilter, is exploited to mitigate potential overfitting in coefficient estimation and enhance the generalization capabilities of the proposed classifier. A series of ablation experiments and parameter analysis were conducted to demonstrate the stability and reliability of the proposed model. Then, we applied DMPNN to 17 publicly available datasets and two engineering applications: Recycling of black plastic wastes and phased resolved partial discharge. The performance results show that the DMPNN model outperforms five classical classifiers and four state-of-the-art (SOTA) class

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Joint segmentation and deformable registration with the presence of the noise using active contour and linear curvature

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AIP Conference Proceedings 2024年第1期3150卷

作者： Nurul Asyiqin Mohd Fauzi Mazlinda Ibrahim Hoo Yann Seong Abdul Kadir Jumaat Lavdie Rada Faculty of Defence Science and Technology National Defence University of Malaysia 57000 Kuala Lumpur Malaysia Centre for Defence Foundation Studies National Defence University of Malaysia 57000 Kuala Lumpur Malaysia School of Mathematical Sciences College of Computing Informatics and Media Universiti Teknologi MARA (UiTM) 40450 Shah Alam Selangor Malaysia Institute for Big Data Analytics and Artificial Intelligence (IBDAAI) Universiti Teknologi MARA (UiTM) 40450 Shah Alam Selangor Malaysia Biomedical Engineering Department Bahcesehir University Besiktas Istanbul Turkey

Medical image processing requires image segmentation and image registration as important diagnostic tools. Image segmentation aims to split structures of interest into distinct components. Meanwhile, image registration finds an ideal transformation between given images. Despite their interrelationship, both tasks are often performed separately. As a result, it faces several problems when undertaken independently. Therefore, the joint models between segmentation and registration have advantages over the separate tasks, particularly when dealing with noisy images. This paper reviewed and compared three existing variational joint segmentation and registration (JSR) models based on active contour without edges and linear curvature. The models are tested on 2D mono-modal and multi-modal images with the presence of white Gaussian noise. The Dice coefficient metric and the registration performance measure are used to evaluate the performance of the models. Numerical results showed that the JSR model using the modified sum of the squared difference outperformed the other two JSR models for noisy mono-modal images. However, the JSR model using the modified normalized gradient fields performed better than the JSR model based on the modified mutual information for noisy multi-modal images.

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Comparison of phase change process in Si-GST hybrid integrated waveguide and MMI devices

arXiv

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arXiv 2020年

作者： Zhang, Hanyu Yang, Xing Lu, Liangjun Chen, Jianping Rahman, B.M.A. Zhou, Linjie College of Telecommunications and Information Engineering Nanjing University of Posts and Telecommunications Nanjing210023 China State Key Laboratory of Advanced Optical Communication Systems and Networks Shanghai Institute for Advanced Communication and Data Science Department of Electronic Engineering Shanghai Jiao Tong University Shanghai200240 China Department of Electrical and Electronic Engineering City University of London LondonEC 1V0HB United Kingdom

In the past decades, silicon photonic integrated circuits (PICs) have been considered as a promising approach to solve the bandwidth bottleneck in optical communications and interconnections. Despite significant advances, large-scale PICs still face a series of technical challenges, such as footprint, power consumption, and routing state storage, resulting from the active tuning methods used to control the optical waves. These challenges can be partially addressed by combining chalcogenide phase change materials (PCMs) such as Ge2Sb2Te5 (GST) with silicon photonics, especially applicable in switching applications due to the nonvolatile nature of the GST. Although GST phase transitions between amorphous and crystalline states actuated by optical and electrical pulses heating have been experimentally demonstrated, there is no direct comparison between them. We carried out simulations and experiments to systematically investigate the difference in the phase change process induced by optical and electrical pulses for two types of Si-GST hybrid waveguides. For the phase transition induced by optical pulses, the device has a clear advantage in terms of power consumption and operation speed. For the phase transition induced by electrical pulses, the device is suitable for large-scale integration because it does not require complex light routing. It helps us better understand the phase change process and push forward the further development of Si-GST hybrid integration platform, bringing in new potential *** Codes 78M50 Copyright © 2020, The Authors. All rights reserved.

关键词： Photonic devices

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Towards Flexible Sparsity-Aware Modeling: Automatic Tensor Rank Learning Using The Generalized Hyperbolic Prior

arXiv

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arXiv 2020年

作者： Cheng, Lei Chen, Zhongtao Shi, Qingjiang Wu, Yik-Chung Theodoridis, Sergios The College of Information Science and Electronic Engineering Zhejiang University Hangzhou310027 China Hangzhou310027 China The Department of Electrical and Electronic Engineering The University of Hong Kong Hong Kong The School of Software Engineering Tongji University Shanghai201804 China The Shenzhen Research Institute of Big Data Shenzhen518172 China The National and Kapodistrian University of Athens Greece Greece The Department of Electronic Systems Aalborg University Denmark

Tensor rank learning for canonical polyadic decomposition (CPD) has long been deemed as an essential yet challenging problem. In particular, since the tensor rank controls the complexity of the CPD model, its inaccurate learning would cause overfitting to noise or underfitting to the signal sources, and even destroy the interpretability of model parameters. However, the optimal determination of a tensor rank is known to be a non-deterministic polynomial-time hard (NP-hard) task. Rather than exhaustively searching for the best tensor rank via trial- and-error experiments, Bayesian inference under the Gaussian-gamma prior was introduced in the context of probabilistic CPD modeling, and it was shown to be an effective strategy for automatic tensor rank determination. This triggered flourishing research on other structured tensor CPDs with automatic tensor rank learning. On the other side of the coin, these research works also reveal that the Gaussian-gamma model does not perform well for high-rank tensors and/or low signal-to-noise ratios (SNRs). To overcome these drawbacks, in this paper, we introduce a more advanced generalized hyperbolic (GH) prior to the probabilistic CPD model, which not only includes the Gaussian-gamma model as a special case, but also is more flexible to adapt to different levels of sparsity. Based on this novel probabilistic model, an algorithm is developed under the framework of variational inference, where each update is obtained in a closed-form. Extensive numerical results, using synthetic data and real-world datasets, demonstrate the significantly improved performance of the proposed method in learning both low as well as high tensor ranks even for low SNR cases. Copyright © 2020, The Authors. All rights reserved.

关键词： Tensors

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The status of the human gene catalogue

arXiv

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arXiv 2023年

作者： Amaral, Paulo Carbonell-Sala, Silvia De La Vega, Francisco M. Faial, Tiago Frankish, Adam Gingeras, Thomas Guigo, Roderic Harrow, Jennifer L. Hatzigeorgiou, Artemis G. Johnson, Rory Murphy, Terence D. Pertea, Mihaela Pruitt, Kim D. Pujar, Shashikant Takahashi, Hazuki Ulitsky, Igor Varabyou, Ales Wells, Christine A. Yandell, Mark Carninci, Piero Salzberg, Steven L. INSPER Institute of Education and Research SP São Paulo Brazil Dr. Aiguader 88 Catalonia Barcelona08003 Spain Department of Biomedical Data Science Stanford University School of Medicine StanfordCA United States Tempus Labs Inc. ChicagoIL United States Nature Genetics San FranciscoCA United States European Molecular Biology Laboratory European Bioinformatics Institute Wellcome Genome Campus Hinxton CambridgeCB10 1SD United Kingdom Department of Functional Genomics Cold Spring Harbor Laboratory Cold Spring HarborNY United States Catalonia Barcelona Spain Centre for Genomics Research Discovery Sciences AstraZeneca Da Vinci Building. Melbourn Science Park RoystonSG8 6HB United Kingdom Universithy of Thessaly Department of Computer Science and Biomedical Informatics Lamia Greece Hellenic Pasteur Institute Athens Greece School of Biology and Environmental Science University College Dublin DublinD04 V1W8 Ireland Conway Institute of Biomedical and Biomolecular Research University College Dublin DublinD04 V1W8 Ireland Department of Medical Oncology Inselspital Bern University Hospital University of Bern Bern3010 Switzerland Department for BioMedical Research University of Bern Bern3008 Switzerland National Center for Biotechnology Information National Library of Medicine National Institutes of Health BethesdaMD20894 United States Center for Computational Biology Johns Hopkins University BaltimoreMD United States Department of Biomedical Engineering Johns Hopkins University BaltimoreMD United States Laboratory for Transcriptome Technology RIKEN Center for Integrative Medical Sciences Kanagawa Yokohama230-0045 Japan Department of Immunology and Regenerative Biology Department of Molecular Neuroscience Weizmann Institute of Science Rehovot76100 Israel Department of Computer Science Johns Hopkins University BaltimoreMD United States Stem Cell Systems Department of Anatomy and Physiology Faculty of Medicine Dentistry and Health Scien

Scientists have been trying to identify all of the genes in the human genome since the initial draft of the genome was published in 2001. Over the intervening years, much progress has been made in identifying protein-coding genes, and the estimated number has shrunk to fewer than 20,000, although the number of distinct protein-coding isoforms has expanded dramatically. The invention of high-throughput RNA sequencing and other technological breakthroughs have led to an explosion in the number of reported non-coding RNA genes, although most of them do not yet have any known function. A combination of recent advances offers a path forward to identifying these functions and towards eventually completing the human gene catalogue. However, much work remains to be done before we have a universal annotation standard that includes all medically significant genes, maintains their relationships with different reference genomes, and describes clinically relevant genetic variants. © 2023, CC BY.

关键词： Genes

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Generating eco-friendly ionic liquids with enhanced CO 2 solubility using language models

Artificial Intelligence Chemistry

引用

Artificial Intelligence Chemistry 2025年第1期3卷

作者： Adroit T.N. Fajar Guillaume Lambard Md. Amirul Islam Bidyut B. Saha Zakiah D. Nurfajrin Kevin Septioga Center for Energy Systems Design (CESD) International Institute for Carbon-Neutral Energy Research (WPI-I2CNER) Kyushu University 744 Motooka Fukuoka 819-0395 Japan Data-driven Materials Design Group Center for Basic Research on Materials National Institute for Materials Science Namiki 1-1 Tsukuba 305-0044 Japan International Institute for Carbon-Neutral Energy Research (WPI-I2CNER) Kyushu University 744 Motooka Fukuoka 819-0395 Japan Department of Applied Chemistry Graduate School of Engineering Kyushu University 744 Motooka Fukuoka 819-0395 Japan

This study presents a viable approach for designing eco-friendly ionic liquids (ILs) with enhanced CO 2 solubility using language models, specifically GPT-2 in conjunction with SMILES-X. The GPT-2 model was fine-tuned on a relatively small, unlabeled IL dataset and subsequently used to generate diverse IL structures. SMILES-X models, trained on IL datasets labeled with CO 2 solubility and eco-toxicity values, were employed to predict the properties of the generated ILs. Trends observed in the predicted IL properties were validated using density functional theory (DFT) and COSMO-RS calculations. The GPT-2 model was then fine-tuned iteratively, with the training data updated by including the top generated ILs from previous cycles. This iterative process led to a gradual improvement in the properties of the generated ILs. It was also observed, however, that continuously adding curated generated ILs to the training data eventually caused the model to produce correct but unrealistic IL structures. These findings highlight both the potential and limitations of language models in designing novel chemicals. Additionally, the CO 2 adsorption capacity of a surrogate IL was experimentally measured, demonstrating the potential of this approach in advancing decarbonization technologies.

关键词： Generative model GPT Prediction SMILES Decarbonization

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Benchmarking in Optimization: Best Practice and Open Issues

arXiv

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arXiv 2020年

作者： Bartz-Beielstein, Thomas Doerr, Carola Bossek, Jakob Chandrasekaran, Sowmya Eftimov, Tome Fischbach, Andreas Kerschke, Pascal López-Ibáñez, Manuel Malan, Katherine M. Moore, Jason H. Naujoks, Boris Orzechowski, Patryk Volz, Vanessa Wagner, Markus Weise, Thomas Institute for Data Science Engineering and Analytics TH Köln Germany Sorbonne Université CNRS LIP6 Paris France Optimisation and Logistics School of Computer Science University of Adelaide Adelaide Australia Computer Systems Department Jožef Stefan Institute Ljubljana Slovenia Statistics and Optimization Group University of Münster Münster Germany School of Computer Science and Engineering University of Málaga Málaga Spain Department of Decision Sciences University of South Africa Pretoria South Africa Institute for Biomedical Informatics University of Pennsylvania PhiladelphiaPA United States Department of Automatics AGH University of Science and Technology Krakow Poland modl.ai Copenhagen Denmark Institute of Applied Optimization School of Artificial Intelligence and Big Data Hefei University Hefei China

This survey compiles ideas and recommendations from more than a dozen researchers with different backgrounds and from different institutes around the world. Promoting best practice in benchmarking is its main goal. The article discusses eight essential topics in benchmarking: clearly stated goals, well-specified problems, suitable algorithms, adequate performance measures, thoughtful analysis, effective and efficient designs, comprehensible presentations, and guaranteed reproducibility. The final goal is to provide well-accepted guidelines (rules) that might be useful for authors and reviewers. As benchmarking in optimization is an active and evolving field of research this manuscript is meant to co-evolve over time by means of periodic *** Codes 68W50. Copyright © 2020, The Authors. All rights reserved.

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Network Alignment

arXiv

引用

arXiv 2025年

作者： Tang, Rui Yong, Ziyun Jiang, Shuyu Chen, Xingshu Liu, Yaofang Zhang, Yi-Cheng Sun, Gui-Quan Wang, Wei School of Cyber Science and Engineering Sichuan University Chengdu610065 China Cyber Science Research Institute Sichuan University Chengdu610065 China Key Laboratory of Data Protection and Intelligent Management Ministry of Education Sichuan University Chengdu610065 China Department of Reproductive Technology The Affiliated Hospital of Southwest Medical University Luzhou646000 China Physics Department University of Fribourg Chemin du Musée 3 Fribourg1700 Switzerland Sino-Europe Complex Science Center School of Mathematics North University of China Shanxi Taiyuan030051 China Complex Systems Research Center Shanxi University Shanxi Taiyuan030006 China School of Public Health Chongqing Medical University Chongqing400016 China

Complex networks are frequently employed to model physical or virtual complex systems. When certain entities exist across multiple systems simultaneously, unveiling their corresponding relationships across the networks becomes crucial. This problem, known as network alignment, holds significant importance. It enhances our understanding of complex system structures and behaviours, facilitates the validation and extension of theoretical physics research about studying complex systems, and fosters diverse practical applications across various fields. However, due to variations in the structure, characteristics, and properties of complex networks across different fields, the study of network alignment is often isolated within each domain, with even the terminologies and concepts lacking uniformity. This review comprehensively summarizes the latest advance- © 2025, CC BY-NC-ND.

关键词： Large scale systems

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Applications of Game Theory in Vehicular Networks: A Survey

arXiv

引用

arXiv 2021年

作者： Sun, Zemin Liu, Yanheng Wang, Jian Li, Guofa Anil, Carie Li, Keqiang Guo, Xinyu Sun, Geng Tian, Daxin Cao, Dongpu College of Computer Science and Technology Jilin University Changchun130012 China Key Laboratory of Symbolic Computation and Knowledge Engineering of Ministry of Education Jilin University Changchun130012 China Institute of Human Factors and Ergonomics College of Mechatronics and Control Engineering Shenzhen University Guangdong Shenzhen518060 China School of Computer Science and Engineering VIT-AP Andhra Pradesh Amaravati522237 India College of Engineering Nanjing Agricultural University Nanjing210031 China State Key Laboratory of Automotive Safety and Energy Department of Automotive Engineering Tsinghua University Beijing100084 China Beijing Advanced Innovation Center for Big Data and Brain Computing Beijing Key Laboratory for Cooperative Vehicle Infrastructure Systems and Safety Control School of Transportation Science and Engineering Beihang University Beijing100191 China Waterloo Cognitive Autonomous Driving Lab University of Waterloo N2L 3G1 Canada

In the Internet of things (IoT) era, vehicles and other intelligent components in an intelligent transportation system (ITS) are connected, forming vehicular networks (VNs) that provide efficient and safe traffic and ubiquitous access to various applications. However, as the number of nodes in an ITS increases, it is challenging to satisfy a varied and large number of service requests with different quality of service (QoS) and security requirements in highly dynamic VNs. Intelligent nodes in VNs can compete or cooperate for limited network resources to achieve the objective for either an individual or a group. Game theory (GT), a theoretical framework designed for strategic interactions among rational decision makers sharing scarce resources, can be used to model and analyze individual or group behaviors of communicating entities in VNs. This paper primarily surveys the recent developments of GT in solving various challenges of VNs. This survey starts with an introduction to the background of VNs. A review of GT models studied in the VNs is then introduced, including the basic concepts, classifications, and applicable vehicular issues. After discussing the requirements of VNs and the motivation of using GT, a comprehensive literature review on GT applications in dealing with the challenges of current VNs is provided. Furthermore, recent contributions of GT to VNs that are integrated with diverse emerging 5G technologies are surveyed. Finally, the lessons learned are given, and several key research challenges and possible solutions of applying GT in VNs are outlined. Copyright © 2021, The Authors. All rights reserved.

关键词： Quality of service

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